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Speciation (7/14) -- Environmental Biology

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Speciation

Speciation Speciation What are species anyway, and how do new ones evolve? Here, you can explore different ways to define a species and learn about the various processes through which speciation can occur. This section also addresses the topics of cospeciation — when two lineages split in concert with one another — and modes of speciation that are specific to plants. Let’s start by defining a species. Defining a species A species is often defined as a group of individuals that actually or potentially interbreed in nature. In this sense, a species is the biggest gene pool possible under natural conditions. For example, these happy face spiders look different, but since they can interbreed, they are considered the same species: Theridion grallator. That definition of a species might seem cut and dried, but it is not — in nature, there are lots of places where it is difficult to apply this definition. For example, many bacteria reproduce mainly asexually. The bacterium shown at right is reproducing asexually, by binary fission. The definition of a species as a group of interbreeding individuals cannot be easily applied to organisms that reproduce only or mainly asexually. Also, many plants, and some animals, form hybrids in nature. Hooded crows and carrion crows look different, and largely mate within their own groups — but in some areas, they hybridize. Should they be considered the same species or separate species? If two lineages of oak look quite different, but occasionally form hybrids with each other, should we count them as different species? There are lots of other places where the boundary of a species is blurred. It’s not so surprising that these blurry places exist — after all, the idea of a species is something that we humans invented for our own convenience! Defining speciation Speciation is a lineage-splitting event that produces two or more separate species. Imagine that you are looking at a tip of the tree of life that constitutes a species of fruit fly. Move down the phylogeny to where your fruit fly twig is connected to the rest of the tree. That branching point, and every other branching point on the tree, is a speciation event. At that point genetic changes resulted in two separate fruit fly lineages, where previously there had just been one lineage. But why and how did it happen? The branching points on this partial Drosophila phylogeny represent long past speciation events. Here is one scenario that exemplifies how speciation can happen: - The scene: a population of wild fruit flies minding its own business on several bunches of rotting bananas, cheerfully laying their eggs in the mushy fruit… - Disaster strikes: A hurricane washes the bananas and the immature fruit flies they contain out to sea. The banana bunch eventually washes up on an island off the coast of the mainland. The fruit flies mature and emerge from their slimy nursery onto the lonely island. The two portions of the population, mainland and island, are now too f
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